The
Diversity of The World of Life
Eukaryotes
(Domain Eukaryota)
Miscellaneous Protists
(various species of eukaryotes
of
unclear
relationships)
Representatives
Various
Amoebae: Various
Radiolarians,
Heliozoons,
Forams, Slime
Molds, etc.
(various species)
Various
Flagellates: Euglenids
(Euglena), Trichonympha (Termite Gut Symbionts), Various Parasites (including
Trypanosoma,
Leishmania, Giardia, & Trichomonas), etc. (various species)
Various
Other Protists (various species)
Biology
ENVIRONMENTS
Protists, in general, live in salt- or fresh-water, in soil, or in the bodies of
other animals.
Most slime molds live in the
shade, as on rocks, leaves, or rotting tree trunks.
Most euglenids live in
freshwater.
DESCRIPTION
The
term "Protists" is something of a catch-all for eukaryotes other
than true plants, fungi, or animals. Although some protists (such as
certain red, brown, or green algae) are multicellular and macroscopic,
most protists are unicellular and microscopic,
traditionally classed as either "protozoans" (primitive animals)
or "algae" (primitive plants).
The "organelles" (membrane-bound bodies
within the "cytoplasm") of a typical, unicellular
protist perform the
various life-sustaining functions that are performed by the
"organs" of multicellular organisms: Although protists are
"primitive" in the sense of being an ancient line of life, their
cells are often far more complex than those of us more
"advanced" forms of life.
Typically
in amoebas, the "ectoplasm" (the
dense outer layer of cytoplasm) gives direction to the creeping movement
of the cell.
Often bearing external filaments, the cells of some protists, such as certain
amoebas, are shelled, often beautifully so. These shells sometimes cement living cells together as
colonies, and the shells of dead protists often accumulate on the ocean
floor as a thick ooze.
Slime
molds are essentially macroscopic amoebas. One
type of slime mold is composed of a "plasmodium" -- a slimy, shapeless
mass of naked cytoplasm with many nuclei (there are no cell walls, cell
membranes, or other
divisions). The other type of slime mold is composed of individual,
amoeboid cells that occasionally coalesce together (in response to a chemical
signal they send out).
Euglenids
are unicellular, with a unique "pellicle" -- a protein-rich,
spiral-ridged coat -- but no
cell wall and with one or two propeller-like "flagella".
FEEDING
HABITS
Most protists feed on "plankton" (microscopic
organisms adrift in water), typically by engulfing their prey.
Some
protists live within the bodies of animals.
This "symbiotic" relationship can be either "commensalism"
(in which the "host" species is not harmed by the protists,
such as those feeding on debris within the gut of vertebrates) or
"mutualism" (in which the host species actually benefits from
the presence of the protists, such as the flagellates pre-digesting fibers
of wood within the gut of termites)
or "parasitism" (in which the host species is harmed by the
protists, such as giardia, which often causes diarrhea in hikers drinking water from mountain streams).
Slime
molds are "heterotrophic" (not producing their own food
molecules). Like other, microscopic amoebae, slime molds engulf
their foodstuffs, which are typically dead organisms or non-living
products of other living organisms (Most slime molds are "saprophytic").
Most
euglenids do not feed but, rather, produce their own food,
photosynthetically, capturing light-energy by means of pigments (green
chlorophylls a and b, as in higher plants) embedded in their
"chloroplasts". In
the absence of light or if cell division has outpaced chloroplast
replication, some euglenids can survive quite well as heterotrophs,
consuming organic matter in the environment; and some colorless euglenid
species are solely predatory. Such "phytoflagellates" as
euglenids have been claimed by both botanists and zoologists (or by neither).
A
euglenid ingests its food by means of a special "feeding apparatus"
(a mouth- and gut-like invagination of the outer cell membrane, of various
degrees of complexity). Note that euglenids were probably originally
predatory and only later photosynthetic -- a prehistoric euglenid presumably ingested
but did not digest a green algal cell (with photosynthetic membranes
inside of its cell membrane), which (surrounded by an envelope of membrane
from the ingesting euglenid) thus became the (triple-membraned) euglenid-chloroplast.
MOTION
As is typical amongst animals, most of the
food-energy consumed by heterotrophic protists ends-up being lost as heat, radiated off
into the environment; however, like most animals, most protists do convert
some of
their food-energy into movement.
"Pseudopods"
(typical amoebas) move by means of "pseudopodia" (arm- or tentacle-like
extensions of the cell).
A
slime mold moves about in a creeping, typically amoeboid manner.
"Zooflagellates" (flagellates traditionally classed as
animals) move by means of long, whip- or propeller-like
"flagella" filaments and sometimes also by means of undulating,
membranous "fins".
A
euglenid typically moves about by means of its one or two propeller-like
flagella, but many may also use their pellicle coat to creep along the
bottom (in a unique, "wriggling" process called "metaboly").
DIGESTION
Water
and other materials are typically exchanged through a
cell membrane via simple diffusion or via "passive" or
"active" transport (both forms of transport employing proteins
embedded within the membrane, the active form also requiring the
biochemical expenditure of energy).
RESPIRATION
Gases
dissolved in environmental water are typically exchanged through a
cell membrane via simple diffusion.
CIRCULATION
The
materials within a eukaryotic
cell are continually mixed by the active process of "cytoplasmic
streaming".
The
materials within the plasmodium of a slime mold are moved by simple
diffusion, by active cytoplasmic streaming (readily observable within the
large mass), and perhaps by the amoeboid movement
of the entire body.
EXCRETION
Within
the cell of a euglenid, or of most other freshwater species of protists, one or two "contractile vacuoles" (water-expelling
"vesicles", or sacs)
help prevent the cell from soaking-up too much water and bursting ("osmolysing"):
Like most substances, water tends to flow from where it is more
concentrated (in the typically freshwater environment) to where it is less
concentrated (within the cell itself, whose cytoplasm is
filled with water-soluble substances).
Water
and other materials are typically exchanged through a
cell membrane via simple diffusion or via "passive" or
"active" transport (both forms of transport employing proteins
embedded within the membrane, the active form also requiring the
biochemical expenditure of energy).
COORDINATION
The activities, growth, and development of protists are under "genetic control" (that is,
under the control of the DNA "chemical blueprint" for the cell),
with feedback from such environmental stimuli as
heat, light, or gravity.
In
euglenids, there is an "eyespot", sensing light by means of a yellow "carotene"
pigment (However, because this is a single-celled organism, there is no
nervous system, as in even the simplest eyes of animals).
The
growth and development of colonial forms of protists, such as cellular
slime molds, is influenced not only by genetics but also by "hormones"
(in effect, chemical
messengers).
REPRODUCTION
Most protists can reproduce "asexually" (that is,
each by itself). Typically,
as a protist grows, its organelles are duplicated; and after reaching a
maximum size (beyond which there is not enough contact between the
contents of the cell and its environment for effective exchange of
materials, both nutrients and wastes), the cell replicates its
"chromosomes" (DNA bodies, within the nucleus) and divides by "fission" (pinching itself in two). This
typical form of cell division (in which the cells produced have the same
number of chromosomes as the parent cells) is called "mitosis"
(Compare "meiosis", in sexual reproduction, below).
Flagellates, such as euglenids, reproduce by "longtitudinal
fission" (splitting in half lengthwise) -- compare ciliates.
In various species of shelled protists, the shell may be divided; the shell
may be retained by one of the offspring and re-grown by the other; or the
shell may at some point prohibit cell-division altogether (See the Asexual
Reproduction of diatoms).
Such blood-sucking insects as mosquitoes or tsetse flies
may carry the infective life stages of certain "pathogenic"
(disease-producing) protists between their various "hosts".
Although euglenids have not been observed to do so, many
protists can reproduce "sexually"; that is, a pair of parent
cells each contribute copies of half of their genetic material to an offspring
-- sexual reproduction, in general, spreads beneficial traits and
"dilutes" harmful traits throughout a population. In
preparation for sexual reproduction, cells divide by "meiosis",
which cuts in half the number of chromosomes in the resultant cells,
typically called male and female "gametes" (sex cells); the
eventual union of gametes produces a "zygote" (a sexually
fertilized egg cell), with both sets of
chromosomes.
Slime
molds occasionally reproduce asexually, by simple fission (dividing in
two). Slime molds also reproduce sexually, in much the same way as fungi,
with which they were traditionally classified (and only recently
distinguished, as by genetic research). Typically, a plasmodium will
produce "sporangia", bodies in which "spores" form, by meiosis (each spore is thus "haploid",
with just half as many chromosomes as in the parent nuclei). The
spores have cell walls (somewhat like those of fungi), which presumably
provide protection during periods of unfavorable environmental
conditions. Eventually, amoeboid or flagellated bodies of naked cytoplasm emerge from
the spores. When a pair of these haploid bodies eventually fuse, they restore the "diploid"
condition (with both sets of chromosomes) in a "zygote" (a
fertilized "egg", so to speak), which either alone or fusing
with others grows into new plasmodium.
Eukaryotes
(Domain Eukaryota)
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