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The Diversity of

The World of Life

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The Diversity of The World of Life

Animals (Metazoa)

Roundworms

(Nematoda)

Representatives

Roundworms, or Nematodes, including elephantiasis parasite, Guinea worm, heartworms, hookworms, Loa loa (African eye worm), pinworms, root-knot nematodes, Trichinella (trichinosis parasite), etc.

Biology

APPROXIMATE NUMBER OF KNOWN SPECIES WORLDWIDE

12,000

ENVIRONMENTS

Roundworms live in saltwater or freshwater, on or in moist soil, or on or in plants or animals.

Nematodes are perhaps the second-most abundant animals on Earth (second only to the arthropods in terms of the number of individuals) -- roundworms outnumber us human beings perhaps a billion-to-one and live on and in almost every living and nonliving thing on the planet (including you and me)!  Fortunately, most are harmless or actually helpful to us (as by feeding on bacteria or pest nematodes).

DESCRIPTION

"Hydrostatic" support (water-pressure) -- within the "pseudocoel" (the unlined body cavity outside of the gut) -- as well as the tough, hard-to-stretch, yet flexible outer "cuticle" give shape to the elongate, cylindrical body, tapering towards both ends.  The cuticle is often sculptured, ringed, or spined; and although it grows in the adult (unlike the exoskeleton of arthropods), it is periodically molted in the young (like the exoskeleton of arthropods).  There may be an adhesive gland at the tail-end.  Nematodes are typically microscopic to minute, but certain species parasitic within vertebrates may grow up to several meters long!

FEEDING HABITS

Roundworms are scavenging, carnivorous, or parasitic (on or in plants or animals).

MOTION

Roundworms have muscles running longitudinally (lengthwise) within the body wall.  Most roundworms move with a characteristic side-to-side, whip-like motion.

DIGESTION

The gut is "complete" -- it has both a mouth and an anus.  The mouth is sunken within a "buccal cavity" (surrounded by such structures as "lips" and bristles) and is often equipped with "teeth" or a piercing/sucking "stylet".

Behind the mouth there is a muscular, food-pumping "pharynx" (sometimes called an "esophagus"), with a one-way valve at its rear end, to prevent regurgitation.  Beyond the pharynx, the gut consists of an "intestine" (in which food materials are digested and from which food materials are absorbed), ending in a "rectum" (which forms the feces).

RESPIRATION

Gases diffuse by osmosis (through selective cell membranes) across the body wall -- there is no respiratory system within these typically thin creatures, as there is in higher animals.

CIRCULATION

The pseudocoel (the unlined body cavity outside of the gut) carries materials circulating throughout the typically small body -- there is no developed circulatory system, as in higher animals.

EXCRETION

"Protonephridia" (as in flatworms) or other primitive excretory systems composed of small internal tubes, "driver" cells, and external openings help maintain the body's salt- and water- balance -- there are no true kidneys.

COORDINATION

The growth, development, and activities of roundworms are under genetic, hormonal, and/or nervous control.

A "nerve ring" and "nerve cords" process nervous-system inputs and outputs.  Although a roundworm has a head, it is poorly "cephalized" -- there are not a great many sensory organs in the head (In particular, "endoparasites", which live within the body of their hosts, typically have little need for elaborate navigation systems).

REPRODUCTION

Most roundworms reproduce sexually.  The reproductive organs typically take up much of the space within the "pseudocoel" (the unlined body cavity outside of the gut) -- the reproductive organs of the female are typically paired, one set in the front half of the body, the other in the rear.  The sexes of roundworms are almost always "separate":  Each individual has either male or female "gonads" -- either "testes" (producing sperms) or "ovaries" (producing eggs) -- but not both.  "Copulation" (mating with physical contact) takes place in many species of roundworms.  Typically, the male uses his curled tail to grab onto the middle of the body of the female.  Then, his pair of short, curved "spicules" reach out from his "cloaca" (his rectum, functioning as the outlet to both the digestive and reproductive systems) and pry open her "vagina" (the female "gonopore", sexual opening).  His sperm (without propeller-like "flagella", as in most animal species) -- which have been stored in his "seminal vesicle" -- are then ejaculated by his "vas deferens" out through his cloaca and into her vagina.  The sperms are stored within her "spermatheca".  The eggs are later fertilized within the body of the female as they pass from the ovaries through the spermatheca and via an "oviduct" to her "uterus".  The fertilized eggs are stored temporarily within the uterus, before being released through the gonopore.  The eggs have a thick shell, which is often sculptured (and, thus, useful in identifying species, such as parasites).

Parasitic species typically have one or more "intermediate" hosts for their larvae, which are different from the "primary" hosts for their adults, in their often complex life cycles.

Animals (Metazoa)

Doug@DouglasDrenkow.com

(c) 2004 D.D.  All Rights Reserved.

Photo of Cells:  H.D.A. Lindquist, US EPA